Withius piger (Simon, 1878)

SPECIES |   ACCEPTED

Original combination: Chelifer piger

LSID urn:lsid:wac.nmbe.ch:name:5452f059-dfd0-43eb-b611-7c38658f9113


Taxonomic references

Chelifer piger

Simon, 1878a: 148–149 original description

Chelifer peculiaris L. Koch:

Pickard-Cambridge, 1889: 111 (misidentification)

Withius piger (Simon):

Vachon, 1970: 186–188, figs 1, 3–6
Heurtault, 1971a: 699–703, 707, figs 22–25, 27
Heurtault, 1971b: 1039–1040, 1045–1046, figs 3a–b, 10a, 11, 12a–b
Mahnert, 1975c: 196
Thaler, 1979: 52
Mahnert, 1980a: 259
Jones, 1980c: n/a, map 24
Harvey, 1981b: 245
Pieper, 1981: 4
Callaini, 1983c: 308–310, figs 3d–f
Callaini, 1986e: 254
Harvey, 1985d: 155
Callaini, 1987b: 274
Callaini, 1988b: 56–57
Legg and Jones, 1988: 129–130, figs 33a, 33b(a–i)
Schaefer, 1988: 99
Schawaller and Dashdamirov, 1988: 44, figs 80–81
Muchmore, 1990a: 522, figs. 18.87, 18.89
Weygoldt, 1990: n/a, fig. 8D
Harvey, 1991a: 663–664
Pieper, 1991: 218
Dashdamirov and Schawaller, 1992b: 60
Gärdenfors and Wilander, 1992: 29–30, fig. 3f
Harvey, 1992c: n/a, figs 205–210
Hogue, 1993: 501
Proctor, 1993: 156 (as Withius subpiger [sic])
Judson, 1994a: n/a, figs 9c–d
Gardini, 1995a: 6
Platen, Blick, Bliss, Drogla, Malten, Martens, Sacher and Wunderlich, 1995: 54
Schawaller, 1995b: 1062
Mahnert, 1997: 576
Poinar, Ćurčić and Cokendolpher, 1998: 90
Bowler, Bullock, Cadbury, Gerlach and Hunter, 1999: 49
Gardini, 2000: 114–115
Harvey, 2001b: 691
Delfosse, 2003b: 39
Ceballos, 2004: 429
Harvey, 2004: 443
Mahnert, 2004: 465
Ozimec, 2004: 389
Pinto, Ribeiro, Silveira Jr and Bernardi, 2005: 115–118 (as Whitius [sic] piger)
Stol, 2005: 14, 33, fig. 18, map 26
Tooren, 2005b: 97
Pascual-Villalobos, Baz and Del Estal, 2005: 233
Pascual-Villalobos, 2006: 1083–1086 (as Whitius [sic] piger)
Zaragoza, 2007a: 63
Holmen and Scharff, 2008: 68
Kunt, Bayram, Yağmur and Danısman, 2008: 81
Harvey, 2010a: 336
Sirvid, Zhang, Harvey, Rhode, Cook, Bartsch and Staples, 2010: 89
Turienzo, Iorio and Mahnert, 2010: 561, 578
Hernández-Teixidor, Castro-Urgal, Txasko, Macías-Hernández and Oromí, 2011: 85
Mahnert, 2011a: 44
Mahnert, 2011d: 37
Zhao, Zhang, Jia and Zhu, 2011: 36
Córdova-Tabares and Villegas-Guzmán, 2013: 602, 610
Mahnert, 2014: 197–198, figs 74–75
Mahnert, Sharaf and Aldawood, 2014: 392
Baert and Mahnert, 2015: 68‒69, map 25
Villegas-Guzmán, 2015: 78
Legg and Farr-Cox, 2016: n/a, fig. 27
Muster and Blick, 2016: 549, 552
Armas, Alegre B., Barba D., Rodríguez-Cabrera, Alayón G. and Peréz G., 2017: 219
Legg, 2019: 191
Červená, Krajčovičová and Christophoryová, 2020b: 223
Červená, Gardini, Jablonski and Christophoryová, 2021: 21, figs 4A, 6D
Lin, Huang, Liu and Chang, 2022: 10
Nassirkhani, 2022: n/a, figs 2B, 2D
Novák, 2024: 83

Not Atemnus piger (Simon):

Beier, 1930a: 209–210 (misidentification; see Diplotemnus ophthalmicus Redikorzev)

Not Miratemnus piger (Simon):

Beier, 1932b: 610 (misidentification; see Diplotemnus ophthalmicus)
Beier, 1932e: 79–80, fig. 99 (misidentification; see Diplotemnus ophthalmicus)
Roewer, 1937: 285 (misidentification; see Diplotemnus ophthalmicus)

Not Diplotemnus piger (Simon):

Beier, 1946: 568–569 (misidentification; see Diplotemnus ophthalmicus)
Beier, 1957b: 147 (misidentification; see Diplotemnus ophthalmicus)
Verner, 1959: 61–63, figs 1–2 (misidentification; see Diplotemnus ophthalmicus)
Beier, 1963b: 248, fig. 251 (misidentification; see Diplotemnus ophthalmicus)
Beier, 1963g: 272 (misidentification; see Diplotemnus ophthalmicus)
Beier, 1965b: 373 (misidentification; see Diplotemnus ophthalmicus)
Beron, 1968: 104 (misidentification; see Diplotemnus ophthalmicus)
Beier, 1971a: 363 (misidentification; see Diplotemnus ophthalmicus)
Schawaller and Dashdamirov, 1988: 34, figs 56–57 (misidentification; see Diplotemnus ophthalmicus)
Schawaller, 1989: 16–17, figs 38–42 (misidentification; see Diplotemnus ophthalmicus)


Synonyms
Chelifer subruber Simon, 1879 -- synonymised by Heurtault, 1971b: 1037 synonymised by Heurtault, 1971b: 1037
LSID urn:lsid:wac.nmbe.ch:name:7b786de5-81e9-4f2c-baed-c8cf063f414c
Chelifer subruber
Simon, 1879a: 30, plate 18 fig. 7 (synonymised by Heurtault, 1971b: 1037) original description
Simon, 1881b: 12
Cantoni, 1882: 192
Tömösváry, 1883: 203–204, plate 2 figs 12–13
Hansen, 1885a: 542–543
Pickard-Cambridge, 1885: 103
Simon, 1885c: 47
Daday, 1889a: 122, 178
Pickard-Cambridge, 1892: 222–223, plate C figs 14, 14a–b
Kulczyński, 1899: 458
Tullgren, 1899a: 170–171
Simon, 1900b: 593
Ellingsen, 1905d: 2
Nosek, 1905: 120, 153
With, 1906: 155–159, figs 5a–b, plate 3, figs 7a–g
Ellingsen, 1907b: 28
Tullgren, 1907a: 37
Bignotti, 1909: 59
Ellingsen, 1909a: 209
Rossi, 1909: 188–289
Warburton, 1909: 438
Ellingsen, 1912a: 122
Ellingsen, 1912b: 85
Ellingsen, 1914: 10
Daday, 1918: 2
Caporiacco, 1923: 131
Schenkel, 1928: 58
Savory, 1945: 180, 184–186
Chelifer (Chelifer) subruber Simon:
Ellingsen, 1910a: 387
Lessert, 1911: 23–24
Chelifer (Withius) subruber Simon:
Kew, 1911a: 49–50, fig. 12
Standen, 1912: 14
Standen, 1916b: 124–125
Standen, 1917: 28–29
Standen, 1918: 335
Kästner, 1928: 8, figs 29–30
Beier, 1929a: 357
Withius subruber (Simon):
Beier, 1930a: 219
Beier, 1930f: 75
Chamberlin, 1931a: n/a, figs 41i, 52g
Beier, 1932e: 198, fig. 202
Beier, 1932f: 488
Roewer, 1936: n/a, figs 45c–d, 63a, 70q, 82, 144
Roewer, 1937: 308
Vachon, 1938a: 40–43, figs 20a–b, 21–22, 52a
Szent-Ivány, 1941: 89
Turk, 1951: 169
Beier, 1952e: 5
Evans and Browning, 1954: 16
Beier, 1956e: 303
Cloudsley-Thompson, 1958b: 96
Beier, 1961b: 37
Meinertz, 1962: 98–100, figs 54c, 55a–b, 56
Beier, 1963b: 282–283, fig. 284
Beier, 1963h: 156
Beier, 1963i: 286
Beier, 1964b: 375
Hoff, 1964b: 36–40
Meinertz, 1964: 400, map 8
Beier, 1966a: 301
Beier, 1966d: 154
Zangheri, 1966: 532
Beier, 1967c: 319
Beier, 1967f: 313
Cloudsley-Thompson, 1968: 127
Hammen, 1969b: 22
Lazzeroni, 1969a: 337–338
Weygoldt, 1969a: 44, 53, 62, 64, 110, figs 47, 58
Weygoldt, 1969b: 230–234, figs 1–2, 3a–c
Beier, 1970a: 45
Lazzeroni, 1970a: 208
Muchmore, 1971a: 94
Schaller, 1971: 429
Verner, 1971: n/a, fig. 62
Weygoldt, 1971a: 14, fig. 23
Legg, 1972a: 576, figs 1(1, 2b), 3(2)
Beier, 1976a: 24
Legg, 1975a: 66
Legg, 1975d: n/a, fig. 2
Weygoldt, 1975b: 310
Beier, 1976f: 244
Beier, 1977b: 6
Crocker, 1978: 9
Schaller, 1979: n/a, plate 10.III fig. 16
Bacallado, 1984: 110
Cloudsley-Thompson, 1984: 190
Kárpáthegyi, 2007a: 89
Kárpáthegyi, 2007b: 96
Distribution
Distribution table
Albania
Algeria
Australia (New South Wales)
Austria
Azerbaijan
Bermuda
Brazil (Rio Grande do Sul)
Chad
Chile
China (Guangdong)
Croatia
Cuba
Denmark
Ecuador (Galapagos Islands)
Egypt
France (mainland)
Germany
Ghana
India
Italy (mainland, Sardinia, Sicily)
Libya
Malta
Mexico
Morocco
Netherlands
New Zealand
Philippines
Portugal (Madeira)
Samoa
Saudi Arabia
Senegal
Seychelles
Solomon Islands
Spain (mainland, Islas Baleares, Islas Canarias)
Sri Lanka
St. Helena
Switzerland
Syria
Taiwan
Tanzania
Tunisia
Turkey
UK
US (Arizona, California, Florida, Georgia, South Carolina)
Type locality: Bou Saâda (as Bou-Saada), M'Sila, Algeria.

Coordinates

Locality Lat Lon
1 Bou Saâda 35.216666 4.183333
Type repository
  • undefined Holotype female (MNHN)
Zoobank LSID

urn:lsid:zoobank.org:act:9D3E8E9B-59FD-4E07-A44A-9AA2C0D8BF22

References

Bacallado, J.J. (1984). Aracnidos. In: Bacallado, J.J. (ed.) Fauna (marina y terrestre) del archipielago Canario Cedirca, Las Palmas de Gran Canaria: 108–115. -- Show included taxa

Beier, M. (1929a). Die Pseudoskorpione des Wiener Naturhistorischen Museums. II. Panctenodactyli. Annalen des Naturhistorischen Museums in Wien 43: 341–367. -- Show included taxa

Beier, M. (1930a). Die Pseudoskorpione des Wiener Naturhistorischen Museums. III. Annalen des Naturhistorischen Museums in Wien 44: 199–222. -- Show included taxa

Beier, M. (1930f). Pseudoscorpione aus Marocco nebst einer Art von der Insel Senafir. Bulletin de la Société des Sciences Naturelles du Maroc 10: 70–77. -- Show included taxa

Beier, M. (1932b). Revision der Atemnidae (Pseudoscorpionidea). Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere 62: 547–610. -- Show included taxa

Beier, M. (1932e). Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich 58: i–xxi, 1–294. -- Show included taxa

Beier, M. (1932f). Pseudoscorpionidea. Spedizione scientifica all'Oasi di Cufra (Marzo-Lugio 1931). Annali del Museo Civico di Storia Naturale di Genova 55: 487–489. -- Show included taxa

Beier, M. (1946). Some pseudoscorpions from the Upper Nile Territory. Annals and Magazine of Natural History (11) 13: 567–571. -- Show included taxa

Beier, M. (1952e). Ordn.: Pseudoscorpionidea, Afterskorpione. In: Strouhal, H. (ed.) Catalogus faunae Austriae Springer-Verlag, Wien 9a: 2–6. -- Show included taxa

Beier, M. (1956e). Ueber Pseudoscorpione aus Spanisch-Marocco. Eos, Madrid 31: 303–310. -- Show included taxa

Beier, M. (1957b). Pseudoscorpione gesammelt von Dr. K. Lindberg 1956. Förhandlingar vid Kungliga Fysiografiska Sällskapets i Lund 27: 145–151. -- Show included taxa

Beier, M. (1961b). Nochmals über iberische und marokkanische Pseudoscorpione. Eos, Madrid 37: 21–39. -- Show included taxa

Beier, M. (1963b). Ordnung Pseudoscorpionidea (Afterskorpione). In: Bestimmungsbücher zur Bodenfauna Europas Akademie-Verlag, Berlin 1: vi, 313 pp.. -- Show included taxa

Beier, M. (1963g). Pseudoskorpione aus Anatolien. Annalen des Naturhistorischen Museums in Wien 66: 267–277. -- Show included taxa

Beier, M. (1963h). Pseudoscorpione aus dem Museum "Enrico Caffi" in Bergamo. Rendiconti dell'Instituto Lombardo di Scienze e Lettere 97B: 147–156. -- Show included taxa

Beier, M. (1963i). Appenninische Pseudoscorpione. Memorie del Museo Civico di Storia Naturale di Verona 10: 283–286. -- Show included taxa

Beier, M. (1964b). Die Pseudoscorpioniden-Fauna Chiles. Annalen des Naturhistorischen Museums in Wien 67: 307–375. -- Show included taxa

Beier, M. (1965b). Pseudoskorpione aus dem Tschad-Gebiet. Annalen des Naturhistorischen Museums in Wien 68: 365–374. -- Show included taxa

Beier, M. (1966a). On the Pseudoscorpionidea of Australia. Australian Journal of Zoology 14: 275–303. -- Show included taxa

Beier, M. (1966d). Die Pseudoscorpioniden der Salomon-Inseln. Annalen des Naturhistorischen Museums in Wien 69: 133–159. -- Show included taxa

Beier, M. (1967c). Die Pseudoscorpione der Noona Dan Expedition nach den Philippinen und Bismarck Inseln. Entomologiske Meddelelser 35: 315–324. -- Show included taxa

Beier, M. (1967f). Ergebnisse zoologischer Sammelreisen in der Türkei. Annalen des Naturhistorischen Museums in Wien 70: 301–323. -- Show included taxa

Beier, M. (1970a). Ergänzungen zur Pseudoskorpionidenfauna der Kanaren. Annalen des Naturhistorischen Museums in Wien 74: 45–49. -- Show included taxa

Beier, M. (1971a). Pseudoskorpione aus dem Iran. Annalen des Naturhistorischen Museums in Wien 75: 357–366. -- Show included taxa

Beier, M. (1976a). Die Pseudoscorpione der macaronesischen Inseln. Vieraea, Tenerife 5: 23–32. -- Show included taxa

Beier, M. (1976f). The pseudoscorpions of New Zealand, Norfolk and Lord Howe. New Zealand Journal of Zoology 3: 199–246. -- Show included taxa

Beier, M. (1977b). Pseudoscorpiones. In: La faune terrestre de l'île de Sainte-Hélène IV. Annales du Musée Royal de l'Afrique Centrale, Zoologie (8) 220: 2–11. -- Show included taxa

Beron, P. (1968). [Etudes sur les Pseudoscorpions. I. Sur les espèces du sous-ordre Cheliferinea en Bulgarie]. Bulletin de l'Institut de Zoologie et Musée, Sofia 27: 103–106. -- Show included taxa

Bignotti, G. (1909). Elenco dei Pseudoscorpioni trovati in Italia e loro distribuzione geografica. Atti della Società dei Naturaliste e Matematici, Modena (4) 11: 56–76. -- Show included taxa

Bowler, J., Bullock, I., Cadbury, J., Gerlach, J. & Hunter, J. (1999). The ecology and conservation of the Aride island, Seychelles. Phelsuma 7: 37–55. -- Show included taxa

Callaini, G. (1983c). Notulae Chernetologicae XII. Nuovi reperti sugli pseudoscorpioni della Sardegna. Lavori della Società Italiana di Biogeografia 8: 279–322. -- Show included taxa

Callaini, G. (1986e). Pseudoscorpioni dell'Italia settentrionale nel Museo Civico di Storia Naturale di Verona (Arachnida). Notulae Chernetologicae XIX. Bollettino del Museo Civico di Storia Naturale, Verona 12: 229–255. -- Show included taxa

Callaini, G. (1987b). Su alcune specie di Cheliferidae della regione Mediterranea (Arachnida, Pseudoscorpionida). (Notulae Chernetologicae. XXII). Bollettino del Museo Civico di Storia Naturale, Verona 13: 273–294. -- Show included taxa

Callaini, G. (1988b). Gli Pseudoscorpioni del Marocco. (Notulae Chernetologicae, XXVII). Annali del Museo Civico di Storia Naturale di Genova 87: 31–66. -- Show included taxa

Cantoni, E. (1882). Escursione in Calabria (1877). Chernetidi ed Opilionidi. Bollettino della Societa Entomologica Italiana 14: 191–203. -- Show included taxa

Caporiacco, L. di (1923). Una nuova specie di Chernetide italiano. Bollettino della Societa Entomologica Italiana 55: 131–134. -- Show included taxa

Ceballos, A. (2004). Pseudoscorpionida. In: Bousquets, J.L., Morrone, J.J., Ordóñez, O.Y. and Fernández, I.V. (ed.) Biodiversidad, taxonomía y biogeografía de Artrópodos de México: hacia una síntesis de su conocimiento Facultad de Ciencias, UNAM, México, D.F. 4: 417–429. -- Show included taxa

Chamberlin, J.C. (1931a). The arachnid order Chelonethida. Stanford University Publications, Biological Sciences 7(1): 1–284. -- Show included taxa

Cloudsley-Thompson, J.L. (1958b). The behaviour of false-scorpions. Naturalist, London 866: 96. -- Show included taxa

Cloudsley-Thompson, J.L. (1968). Spiders, scorpions, centipedes and mites. Pergamon Press, Oxford. [revised edition] -- Show included taxa

Cloudsley-Thompson, J.L. (1984). Arachnids. In: Cloudsley-Thompson, J.L. (ed.) Key environments. Sahara Desert Pergamon Press, Oxford: 175–204. -- Show included taxa

Córdova-Tabares, V. & Villegas-Guzmán, G. (2013). Nuevos registros de pseudoescorpiones (Arachnida: Pseudoscorpiones) en Chiapas, México. Acta Zoológica Mexicana (nueva serie) 29: 596–613. -- Show included taxa

Crocker, J. (1978). Introduction to false scorpions. British Arachnological Society, Secretary's News Letter 21: 4–9. -- Show included taxa

Daday, E. (1889a). A Magyar Nemzeti Muzeum álskorpióinak áttekintése. Természetrajzi Füzetek 11: 111–136, 165–192. -- Show included taxa

Daday, E. (1918). Ordo Pseudoscorpiones. In: A Magyar Birodalom Allatvilága Regia Societas Scientiarum Naturalium Hungarica, Budapest: 1–2. -- Show included taxa

Dashdamirov, S. & Schawaller, W. (1992b). [Pseudoscorpions of the Caucasian fauna (Arachnida Pseudoscorpionida)]. Arthropoda Selecta 1(4): 31–72. -- Show included taxa

Delfosse, E. (2003b). Catalogue préliminaire des Pseudoscorpions de France métropolitaine (Arachnida Pseudoscorpiones). Bulletin de Phyllie 17: 24–48. -- Show included taxa

Ellingsen, E. (1905d). Pseudoscorpions from Italy and southern France conserved in the R. Museo Zoologico in Torino. Bollettino dei Musei di Zoologia e di Anatomia Comparata della R. Università di Torino 20(503): 1–13. -- Show included taxa

Ellingsen, E. (1907b). Über einige Pseudoskorpione aus Deutsch-Ostafrika. Zoologischer Anzeiger 32: 28–30. -- Show included taxa

Ellingsen, E. (1909a). Contributions to the knowledge of the pseudoscorpions from material belonging to the Museo Civico in Genova. Annali del Museo Civico di Storia Naturale di Genova (3) 4: 205–220. -- Show included taxa

Ellingsen, E. (1910a). Die Pseudoskorpione des Berliner Museums. Mitteilung aus dem Zoologischen Museum in Berlin 4: 357–423. -- Show included taxa

Ellingsen, E. (1912a). H. Sauter's Formosa-Ausbeute. Pseudoscorpions from Formosa. I. Nyt Magazin for Naturvidenskaberne 50: 121–128. -- Show included taxa

Ellingsen, E. (1912b). The pseudoscorpions of South Africa, based on the collections of the South African Museum, Cape Town. Annals of the South African Museum 10: 75–128. -- Show included taxa

Ellingsen, E. (1914). On the pseudoscorpions of the Indian Museum, Calcutta. Records of the Indian Museum 10: 1–14. -- Show included taxa

Evans, G.O. & Browning, E. (1954). Synopses of the British Fauna. No. 10. Pseudoscorpiones. Linnean Society, London. -- Show included taxa

Gärdenfors, U. & Wilander, P. (1992). Sveriges klokrypare med nyckel till arterna. Entomologisk Tidskrift 113: 20–35. -- Show included taxa

Gardini, G. (1995a). Arachnida Pseudoscorpionida. In: Minelli, A, Ruffo, S. and La Posta, S. (ed.) Checklist delle specie della fauna Italiana Calderini, Bologna 22: 1–8. -- Show included taxa

Gardini, G. (2000). Catalogo degli Pseudoscorpioni d'Italia (Arachnida). Fragmenta Entomologica 32, Supplemento: 1–181. -- Show included taxa

Hammen, L. van der (1969b). Bijdrage tot de kennis van de Nederlandse bastaardschorpioenen (Arachnida, Pseudoscorpionida). Zoologische Bijdragen 11: 15–24. -- Show included taxa

Hansen, H.J. (1885a). Arthrogastra Danica: en monographisk fremstilling af de i Danmark levende Meiere og Mosskorpioner med bidrag til sidstnaevnte underordens systematik. In: Naturhistorisk Tidsskrift Thieles Bogtrykkeri, Kjøbenhavn (3) 14: 491–554. -- Show included taxa

Harvey, M.S. (1981b). A checklist of the Australian Pseudoscorpionida. Bulletin of the British Arachnological Society 5: 237–252. -- Show included taxa

Harvey, M.S. (1985d). Pseudoscorpionida. In: Walton, D.W. (ed.) Zoological catalogue of Australia Australian Government Publishing Service, Canberra 3: 126–155. -- Show included taxa

Harvey, M.S. (1991a). Catalogue of the Pseudoscorpionida. Manchester University Press, Manchester. -- Show included taxa

Harvey, M.S. (1992c). The phylogeny and classification of the Pseudoscorpionida (Chelicerata: Arachnida). Invertebrate Taxonomy 6: 1373–1435. -- Show included taxa

Harvey, M.S. (2001b). The Schizomida (Arachnida) of the Seychelle Islands. Invertebrate Taxonomy 15: 681–693. doi: https://doi.org/10.1071/IT01006 -- Show included taxa

Harvey, M.S. (2004). Remarks on the New World pseudoscorpion genera Parawithius and Victorwithius, with a new genus bearing a remarkable sternal modification (Pseudoscorpiones, Withiidae). Journal of Arachnology 32: 436–456. -- Show included taxa

Harvey, M.S. (2010a). A synopsis of the smaller arachnid orders of the Seychelles islands (Arachnida: Pseudoscorpiones, Schizomida, Amblypygi and Palpigradi). In: Gerlach, J. and Marusik, Y. (ed.) Arachnida and Myriapoda of the Seychelles Islands Siri Scientific Press, Manchester: 331–340. -- Show included taxa

Hernández-Teixidor, D., Castro-Urgal, R., Txasko, N., Macías-Hernández, N. & Oromí, P. (2011). Fauna de arácnidos del malpaís de La Rasca (Tenerife, Islas Canarias). Vieraea 39: 77–95. -- Show included taxa

Heurtault, J. (1971a). Pseudoscorpions de la région du Tibesti (Sahara méridionale). IV. Cheliferidae. Bulletin du Muséum National d'Histoire Naturelle, Paris (2) 42: 685–707. -- Show included taxa

Heurtault, J. (1971b). Chambre génitale, armature génitale et caractères sexuels secondaires chez quelques espèces de Pseudoscorpions (Arachnida) du genre Withius. Bulletin du Muséum National d'Histoire Naturelle, Paris (2) 42: 1037–1053. -- Show included taxa

Hoff, C.C. (1964b). Atemnid and cheliferid pseudoscorpions, chiefly from Florida. American Museum Novitates 2198: 1–43. -- Show included taxa

Hogue, C.L. (1993). Latin American insects and entomology. University of California Press, Berkeley. -- Show included taxa

Holmen, M. & Scharff, N. (2008). Anthrenochernes stellae Lohmander, 1939 - status in Denmark for a new species on the EC Habitats Directive (Arachnida, Pseudoscorpiones). Entomologiske Meddelelser- Entomologisk Forening Kobenhavn 76: 55–68. -- Show included taxa

Jones, P.E. (1980c). Provisional atlas of the Arachnida of the British Isles. Part 1. Pseudoscorpiones.. Institute of Terrestrial Ecology, Monks Wood. -- Show included taxa

Judson, M.L.I. (1994a). Les Pseudoscorpions. Pénélope 12: 1–37. -- Show included taxa

Kárpáthegyi, P. (2007a). Pseudoscorpions of Hungary. Folia Historico Naturalia Musei Matraensis 31: 81–90. -- Show included taxa

Kárpáthegyi, P. (2007b). Check list of the Hungarian Pseudoscorpiones. Folia Historico Naturalia Musei Matraensis 31: 91–97. -- Show included taxa

Kästner, A. (1928). 2. Ordnung: Moos- oder Afterskorpione, Pseudoscorpiónes Latr. (Chernétes Simon; Chelonéti Thorell; Chernetídea Camb.). In: Brohmer, P, Ehrmann, P. and Ulmer, G. (ed.) Die Tierwelt Mitteleuropas Quelle und Meyer, Leipzig 3 (1, IV): 1–13. -- Show included taxa

Kew, H.W. (1911a). A synopsis of the false scorpions of Britain and Ireland. Proceedings of the Royal Irish Academy, section B 29: 38–64. -- Show included taxa

Kulczyński, V. (1899). Arachnoidea opera Rev. E. Schmitz collecta insulis Maderianis et in insulis Selvages dictis. Rozpravy Akademia Umiejetnosei 36: 319–461. -- Show included taxa

Kunt, K.B., Bayram, A., Yağmur, E.A. & Danısman, T. (2008). Checklist of the pseudoscorpions of Turkey (Pseudoscorpionida; Arachnida). Turkish Journal of Arachnology 1: 70–84. -- Show included taxa

Lazzeroni, G. (1969a). Sur la faune de pseudoscorpions de la région apenninique méridionale. (Recherches sur les Pseudoscorpions. III.). Memorie del Museo Civico di Storia Naturale di Verona 16: 321–344. -- Show included taxa

Lazzeroni, G. (1970a). Ricerche sugli Pseudoscorpioni. III. Considerazioni biogeografiche sulla fauna della regione appenninica meridionale. Bulletin du Muséum National d'Histoire Naturelle, Paris (2) 41(supplément 1): 205–208. -- Show included taxa

Legg, G. (1972a). False-scorpions: the families Cheliferiidae [sic] and Chernetiidae [sic]. Countryside 21: 576–583. -- Show included taxa

Legg, G. (1975a). A generalised account of the male genitalia and associated glands of pseudoscorpions (Arachnida). Bulletin of the British Arachnological Society 3: 66–74. -- Show included taxa

Legg, G. (1975d). The possible significance of spermathecae in pseudoscorpions (Arachnida). Bulletin of the British Arachnological Society 3: 91–95. -- Show included taxa

Legg, G. & Jones, R.E. (1988). Synopses of the British fauna (new series). 40. Pseudoscorpions (Arthropoda; Arachnida). Brill/Backhuys, Leiden. -- Show included taxa

Lessert, R. de (1911). Pseudoscorpions. Catalogue des Invertébrés de la Suisse 5: 1–50. -- Show included taxa

Mahnert, V. (1975c). Pseudoscorpione von den maltesischen Inseln. Fragmenta Entomologica 11: 185–197. -- Show included taxa

Mahnert, V. (1980a). Pseudoscorpions from the Canary Islands. Entomologica Scandinavica 11: 259–264. -- Show included taxa

Mahnert, V. (1997). New species and records of pseudoscorpions (Arachnida, Pseudoscorpiones) from the Canary Islands. Revue Suisse de Zoologie 104: 559–585. -- Show included taxa

Mahnert, V. (2004). Die Pseudoskorpione Österreichs (Arachnida, Pseudoscorpiones). Denisia 12: 459–471. -- Show included taxa

Mahnert, V. (2011a). A nature’s treasury: Pseudoscorpion diversity of the Canary Islands, with the description of nine new species (Pseudoscorpiones, Chthoniidae, Cheiridiidae) and new records. Revista Ibérica de Aracnología 19: 27–45. -- Show included taxa

Mahnert, V. (2011d). Pseudoscorpiones (Arachnida). In: Schuster, R. (ed.) Checklisten der Fauna Österreichs Verlag der Österreichischen Akademie der Wissenschaften, Wien 5: 28–39. -- Show included taxa

Mahnert, V. (2014). Pseudoscorpions (Arachnida: Pseudoscorpiones) from the Galapagos Islands (Ecuador). Revue Suisse de Zoologie 121: 135–210. -- Show included taxa

Mahnert, V., Sharaf, M. & Aldawood, A.S. (2014). Further records of pseudoscorpions (Arachnida, Pseudoscorpiones) from Saudi Arabia. Zootaxa 3764: 387–393. -- Show included taxa

Meinertz, T. (1962). Mosskorpioner og mejere (Pseudoscorpionidea og Opiliones). G.E.C. Gads, København 67. -- Show included taxa

Meinertz, T. (1964). Beiträge zur Verbreitung der Pseudoskorpioniden in Dänemark. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn 126: 387–402. -- Show included taxa

Muchmore, W.B. (1971a). Phoresy by North and Central American pseudoscorpions. Proceedings of the Rochester Academy of Science 12: 79–97. -- Show included taxa

Muchmore, W.B. (1990a). Pseudoscorpionida. In: Dindal, D.L. (ed.) Soil biology guide John Wiley and Sons, New York: 503–527. -- Show included taxa

Nosek, A. (1905). Araneiden, Opilionen und Chernetiden. In: Penther, A., Zederbauer, E. Ergebnisse einer naturwissenschaftliche Reise zum Erdschais-Dagh (Kleinasien). Annalen des K. K. Naturhistorischen Hofmuseums 20: 114–154. -- Show included taxa

Ozimec, R. (2004). List of Croatian pseudoscorpion fauna (Arachnida, Pseudoscorpiones). Natura Croatica 13: 381–394. -- Show included taxa

Pascual-Villalobos, M.J. (2006). Psocids: weight losses of grain and biological control by pseudoscorpions. In: Lorini, I., Bacaltchuk, B., Beckel, H., Deckers, D., Sundfeld, E., dos Santos, J.P., Biagi, J.D., Celaro, J.C., Faroni, L.R.D'A., Bortolini, L.de O.F., Sartori, M.R., Elias, M.C., Guedes, R.N.C., Fonseca, R.G. da and Scussel, V.M. (ed.) Proceedings of the 9th International Working Conference on Stored Product Protection, 15 to 18 October 2006, Campinas, São Paulo, Brazil. Brazilian Post-harvest Association - ABRAPOS, Passo Fundo, RS, Brazil, 2006: 1083–1086. -- Show included taxa

Pascual-Villalobos, M.T., Baz, A. & del Estal, P. (2005). Occurrence of psocids and natural predators on organic rice in Calasparra (Murcia, Spain). Journal of Stored Products Research 41: 231–235. -- Show included taxa

Pickard-Cambridge, O. (1885). Pseudoscorpions new to Britain. Naturalist, London 10: 103. -- Show included taxa

Pickard-Cambridge, O. (1889). On new and rare British spiders. Proceedings of the Dorset Natural History and Antiquarian Field Club and Archaeological Society 10: 107–138. -- Show included taxa

Pickard-Cambridge, O. (1892). On the British species of false-scorpions. Proceedings of the Dorset Natural History and Antiquarian Field Club and Archaeological Society 13: 199–231. -- Show included taxa

Pieper, H. (1981). Die Pseudoskorpione von Madeira und nachbarinseln. Bocagiana, Funchal 60: 1–7. -- Show included taxa

Pieper, H. (1991). New records of pseudoscorpions from the Madeiran Islands. Boletim do Museu Municipal do Funchal 42: 217–221. -- Show included taxa

Pinto, D.M., Ribeiro, P.B., Silveira Jr, P. & Bernardi, E. (2005). Population flotation of Whitius [sic] piger (Pseudoscorpionida) in poultry farm in Pelotas, RS, Brazil. Archives of Veterinary Science 10: 115–118. -- Show included taxa

Platen, R., Blick, T., Bliss, P., Drogla, R., Malten, A., Martens, J., Sacher, P. & Wunderlich, J. (1995). Verzeichnis der Spinnentiere (excl. Acarida) Deutschlands (Arachnida: Araneida, Opilionida, Pseudoscorpionida). Arachnologische Mitteilungen, Sonderband 1: 1–55. -- Show included taxa

Poinar, G.O., Jr, Ćurčić, B.P.M. & Cokendolpher, J.C. (1998). Arthropod phoresy involving pseudoscorpions in the past and present. Acta Arachnologica 47: 79–96. -- Show included taxa

Proctor, H.C. (1993). Mating biology resolves trichotomy for cheliferoid pseudoscorpions (Pseudoscorpionida, Cheliferoidea). Journal of Arachnology 21: 156–158. -- Show included taxa

Roewer, C.F. (1936). Chelonethi oder Pseudoskorpione. In: Bronns, H.G. (ed.) Bronn's Klassen und Ordnungen des Tierreichs Akademische Verlagsgesellschaft M.B.H., Leipzig 5(IV)(6)(1): 1–160. -- Show included taxa

Roewer, C.F. (1937). Chelonethi oder Pseudoskorpione. In: Bronns, H.G. (ed.) Bronn's Klassen und Ordnungen des Tierreichs Akademische Verlagsgesellschaft M.B.H., Leipzig 5(IV)(6)(1): 161–320. -- Show included taxa

Rossi, A. (1909). Materiali per una fauna aracnologica della Provincia di Roma. Chernetes (Pseudoscorpioni). Bollettino della Societa Entomologica Italiana 40: 182–194. -- Show included taxa

Savory, T.H. (1945). The spiders and allied orders of the British Isles comprising descriptions of every family of British spiders, every species of harvestmen and false scorpions also the more familiar of the British mites and sea-spiders. Frederick Warne & Co., London. [2nd edition] -- Show included taxa

Schaefer, M. (1988). 1. Chelicerata, Spinnentiere. In: Brohmer, P. (ed.) Fauna von Deutschland Quelle und Meyer, Heidelberg: 99–135. [17th edition] -- Show included taxa

Schaller, F. (1971). Indirect sperm transfer by soil arthropods. Annual Review of Entomology 16: 407–446. -- Show included taxa

Schaller, F. (1979). Significance of sperm transfer and formation of spermatophores in arthropod phylogeny. In: Gupta, A.P. (ed.) Arthropod phylogeny Van Nostrand Reinhold, New York: 587–608. -- Show included taxa

Schawaller, W. (1989). Pseudoskorpione aus der Sowjetunion, Teil 3 (Arachnida: Pseudoscorpiones). Stuttgarter Beiträge zur Naturkunde (A) 440: 1–30. -- Show included taxa

Schawaller, W. (1995b). Review of the pseudoscorpion fauna of China (Arachnida: Pseudoscorpionida). Revue Suisse de Zoologie 102: 1045–1064. -- Show included taxa

Schawaller, W. & Dashdamirov, S. (1988). Pseudoskorpione aus dem Kaukasus, Teil 2 (Arachnida). Stuttgarter Beiträge zur Naturkunde (A) 415: 1–51. -- Show included taxa

Schenkel, E. (1928). Pseudoscorpionida (Afterskorpione). In: Dahl, F. (ed.) Die Tierwelt Deutschlands G. Fischer, Jena 8: 52–72. -- Show included taxa

Simon, E. (1878a). Liste des espèces de la famille des Cheliferidae qui habitant l'Algérie et le Maroc. Annales de la Société Entomologique de France (5) 8: 144–153. -- Show included taxa

Simon, E. (1879a). Les Ordres des Chernetes, Scorpiones et Opiliones. In: Les Arachnides de France Librairie Encyclopédique de Roret, Paris 7: 1–332. -- Show included taxa

Simon, E. (1881b). Descriptions d'Arachnides nouveaux d'Afrique. (Chernetes de la Basse Égypte rec. par M. Letourneux). Bulletin de la Société Zoologique de France 6: 1–15. -- Show included taxa

Simon, E. (1885c). Étude sur les Arachnides recueillis en Tunisie en 1883 et 1884 par Mm. A. Letourneux, M. Sédillot et Valery Mayet. In: Exploration Scientifique de la Tunisie Imprimerie Nationale, Paris: 1–55. -- Show included taxa

Simon, E. (1900b). Studio sui Chernetes Italiani conservati nel Museo Civico di Genova. II. Annali del Museo Civico di Storia Naturale di Genova (2) 20: 593–595. -- Show included taxa

Sirvid, P.J., Zhang, Z.-Q., Harvey, M.S., Rhode, B.E., Cook, D.R., Bartsch, I. & Staples, D.A. (2010). Phylum Arthropoda, Chelicerata, horseshoe crabs, arachnids and sea spiders. In: Gordon, D. (ed.) The New Zealand inventory of biodiversity: a Species 2000 Symposium review Canterbury University Press, Christchurch 1: 50–89. -- Show included taxa

Standen, R. (1912). The false-scorpions of Lancashire and some adjoining counties, with a preliminary list of records. Lancashire and Cheshire Naturalist 5: 7–16. -- Show included taxa

Standen, R. (1916b). Chelifer (Chernes) powelli Kew and Chelifer (Withius) subruber Simon in Lancashire and Cheshire. Lancashire and Cheshire Naturalist 9: 124–125. -- Show included taxa

Standen, R. (1917). Report on the false-scorpions (Chelifers). Lancashire and Cheshire Naturalist 10: 27–31. -- Show included taxa

Standen, R. (1918). Report on the false-scorpions (Chelifers) for 1917. Lancashire and Cheshire Naturalist 10: 335. -- Show included taxa

Stol, I. (2005). Nordiske mosskorpioner (Pseudoscorpiones). Norske Insekttabeller 18: 1–35. -- Show included taxa

Szent-Ivány, J. (1941). Neue Angaben zur Verbreitung der Pseudoscorpione im Karpaten-becken. Fragmenta Faunistica Hungarica 4: 85–90. -- Show included taxa

Thaler, K. (1979). Fragmenta faunistica Tirolensia, IV (Arachnida: Acari: Caeculidae; Pseudoscorpiones; Scorpiones; Opiliones; Aranei; Insecta: Dermaptera; Thysanoptera; Diptera Nematocera: Mycetophilidae, Psychodidae, Limoniidae und Tipulidae). Veröffentlichungen des Museums Ferdinandeum 59: 49–83. -- Show included taxa

Tömösváry, Ö. (1883). Pseudoscorpiones faunae Hungaricae. A Magyar fauna álskorpiói. Magyar Tudományos Akadémia Matematikai és Természettudományi Közlemények 18: 135–256. -- Show included taxa

Tooren, D. van den (2005b). Naamlijst en determinatiesleutel tot de Pseudoschorpioenen van Nederland (Arachnida: Pseudoscorpiones). Nederlandse Faunistische Mededelingen 23: 91–102. -- Show included taxa

Tullgren, A. (1899a). Bidrag till Kännedomen om Sveriges Pseudoscorpioner. Entomologisk Tidskrift 20: 161–182. -- Show included taxa

Tullgren, A. (1907a). Zur Kenntnis aussereuropäischer Chelonethiden des Naturhistorischen Museums in Hamburg. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 24: 21–75. -- Show included taxa

Turienzo, P., Iorio, O. di & Mahnert, V. (2010). Global checklist of pseudoscorpions (Arachnida) found in birds’ nests. Revue Suisse de Zoologie 117: 557–598. -- Show included taxa

Turk, F.A. (1951). On the swarming of pseudoscorpions and their association with ants. Entomologist's Monthly Magazine 87: 169. -- Show included taxa

Vachon, M. (1938a). Recherches anatomiques et biologiques sur la réproduction et le développement des Pseudoscorpions. Annales des Sciences Naturelles, Zoologie (11) 1: 1–207. -- Show included taxa

Vachon, M. (1970). Remarques sur Withius piger (Simon, 1878) nov. comb. (Pseudoscorpion Cheliferidae) et sur le genre Diplotemnus J.C. Chamberlin, 1933, à propos de Diplotemnus beieri nov. nom. (Pseudoscorpion Miratemnidae). Bulletin du Muséum National d'Histoire Naturelle, Paris (2) 42: 185–191. -- Show included taxa

Verner, P.H. (1959). Ein interessanter Fund eines Pseudoscorpions in der Tschechoslowakei (Pseudoscorpionidea). Acta Faunistica Entomologica Musei Nationalis Pragae 5: 61–63. -- Show included taxa

Verner, P.H. (1971). Štírci - Pseudoscorpionidea. In: Daniel, M. and Černý, V. (ed.) Klič Zviřeny ČSSR Českoslovenká Akademie Věd, Praha 4: 19–31. -- Show included taxa

Villegas-Guzmán, G.A. (2015). Psudoescorpiones (Arachnida: Pseudoscorpiones) de la Ciudad de México y sus alrededores. Entomología Mexicana 2: 76–81. -- Show included taxa

Warburton, C. (1909). Arachnida Embolobranchiata (scorpions, spiders, mites, etc.). In: Harmer, S.F. and Shipley, A.E. (ed.) The Cambridge natural history MacMillan and Co., London 4: 297–474. -- Show included taxa

Weygoldt, P. (1969a). The biology of pseudoscorpions. Harvard University Press, Cambridge, Massachusetts. -- Show included taxa

Weygoldt, P. (1969b). Paarungsverhalten und Samenübertragung beim Pseudoskorpion Withius subruber Simon (Cheliferidae). Zeitschrift für Tierpsychologie 26: 230–235. -- Show included taxa

Weygoldt, P. (1971a). Vergleichend-embryologische Untersuchungen an Pseudoscorpionen V. Das Embryonalstadium mit seinem Pumporgan bei verschiedenen Arten und sein Wert als taxonomisches Merkmal.. Zeitschrift für die Zoologische Systematik und Evolutionforschung 9: 3–29. -- Show included taxa

Weygoldt, P. (1975b). Die indirekte Spermatophorenübertragung bei Arachniden. Verhandlungen der Deutschen Zoologischen Gesellschaft 1974: 308–313. -- Show included taxa

Weygoldt, P. (1990). Arthropoda-Chelicerata: sperm transfer. In: Adiyodi, K.G. and Adiyodi, R.G. (ed.) Reproductive biology of invertebrates Oxford & IBH Publishing Co., New Delhi 4 (B): 77–119. -- Show included taxa

With, C.J. (1906). The Danish expedition to Siam 1899–1900. III. Chelonethi. An account of the Indian false-scorpions together with studies on the anatomy and classification of the order. Oversigt over det Konigelige Danske Videnskabernes Selskabs Forhandlinger (7) 3: 1–214. -- Show included taxa

Zangheri, P. (1966). Repertorio sistematico e topografico della flora e fauna vivente e fossile della Romagna. Museo Civico di Storia Naturale di Verona, Memorie Fuori Serie 1 2: 485–854. -- Show included taxa

Zaragoza, J.A. (2007a). Catálogo de los Pseudoescorpiones de la Península Ibérica e Islas Baleares (Arachnida: Pseudoscorpiones). Revista Ibérica de Aracnología 13: 3–91. -- Show included taxa

Zhao, Y.-W., Zhang, F., Jia, Y. & Zhu, M.-S. (2011). [A checklist of pseudoscorpions from China (Arachnida: Pseudoscorpiones)]. Acta Arachnologica Sinica 20: 3041. -- Show included taxa

Armas, L.F. de, Alegre B., A., Barba D., R.A., Rodríguez-Cabrera, T.M., Alayón G., G. & Peréz G., A. (2017). Arácnidos. In: C. A. Mancina and D. D. Cruz F. Diversidad biológica de Cuba: métodos de inventario, monitoreo y colecciones biológicas Editorial AMA: La Habana: 196–223. -- Show included taxa

Legg, G. (2019). Changes in the British pseudoscorpion fauna over the last 50 years. Arachnology 18: 189–195. -- Show included taxa

Lin, H.Y., Huang, J.X., Liu, H.H. & Chang, C.-H. (2022). Two new pseudoscorpion species of the coastal genus Garypus L. Koch, 1873 (Garypidae) and an updated checklist of the Pseudoscorpiones of Taiwan. Zoological Studies 61(24): 1–13. -- Show included taxa

Červená, M., Gardini, G., Jablonski, D. & Christophoryová, J. (2021). Checklist of pseudoscorpions (Arachnida, Pseudoscorpiones) of Albania. Zoological Studies 60(17): 1–25. -- Show included taxa

Nassirkhani, M. (2022). Description of a new pseudoscorpion, Withius leggi sp. n. (Pseudoscorpiones: Withiidae) from northern Iran. Zoology in the Middle East 68: 272–279. -- Show included taxa

Legg, G. & Farr-Cox, F. (2016). Illustrated key to the British false scorpions (pseudoscorpions). Field Studies Council: Telford. -- Show included taxa

Baert, L. & Mahnert, V. (2015). The distribution of the non‐Araneae and non‐Acari arachnids of Galápagos. Belgian Journal of Entomology 28: 1–76. -- Show included taxa

Muster, C. & Blick, T. (2016). Rote Liste und Gesamtartenliste der Pseudoskorpione (Arachnida: Pseudoscorpiones) Deutschlands.. Naturschutz und Biologische Vielfalt 70: 539–561. -- Show included taxa

Novák, J. (2024). An updated checklist for the pseudoscorpion fauna of Hungary (Arachnida: Pseudoscorpiones). Zootaxa 5433: 51–95. -- Show included taxa

Červená, M., Krajčovičová, K. & Christophoryová, J. (2020b). Updated checklist of pseudoscorpions (Arachnida: Pseudoscorpiones) of Central Europe. Arthropoda Selecta 29: 219–228.. Arthropoda Selecta 29: 219–228. -- Show included taxa

Logs
Date Type Reference Old value
2024-09-28 Taxonomic reference update Červená, Krajčovičová and Christophoryová, 2020
2024-09-28 New taxonomic reference entry Novák, 2024
2024-09-28 New taxonomic reference entry Červená, Krajčovičová and Christophoryová, 2020
2024-09-28 Species updated Old value
2023-04-22 New taxonomic reference entry Muster and Blick, 2016
2023-01-22 Species synonymy Heurtault, 1971
2023-01-22 Taxonomic reference update Pinto, Ribeiro, Silveira Jr and Bernardi, 2005
2023-01-22 Taxonomic reference update Pascual-Villalobos, 2006
2022-10-30 New taxonomic reference entry Baert and Mahnert, 2015
2022-10-03 New taxonomic reference entry Legg and Farr-Cox, 2016
2022-09-26 New taxonomic reference entry Nassirkhani, 2022
2022-09-26 Taxonomic reference update Lin, Huang, Liu and Chang, 2022
2022-09-26 Species updated Old value
2022-09-26 New taxonomic reference entry Lin, Huang, Liu and Chang, 2022
2022-09-26 New taxonomic reference entry Červená, Gardini, Jablonski and Christophoryová, 2021
2022-09-26 New taxonomic reference entry Legg, 2019
2022-03-05 New taxonomic reference entry Armas, Alegre B., Barba D., Rodríguez-Cabrera, Alayón G. and Peréz G., 2017